1. Faculty of Agriculture, Ehime University, Matsuyama, Ehime 790-8566, Japan
2. Department of Agronomy, Kansas State University, Manhattan, KS 66506, U.S.A.
3. National Institute of Agrobiological Sciences, Tsukuba, Ibaraki 305-8602, Japan
4. Hard Winter Wheat Genetics Research Unit, U.S. Department of Agriculture-Agriculture Research Service, Manhattan, KS 66506, U.S.A.