1. Caesar Kleberg Wildlife Research Institute, Texas A&M University-Kingsville, Kingsville, TX 78363, USA
2. Department of Biological Sciences, Western Michigan University, 1903 W Michigan Avenue, Kalamazoo, MI 49008, USA
3. Department of Biology, Grand Valley State University, 1 Campus Drive, Allendale, MI 49401, USA
4. Department of Ecology and Evolutionary Biology, University of Tennessee, 1502 Cumberland Avenue, Knoxville, TN 37996, USA
5. Department of Human Genetics, University of Michigan, 500 S. State Street, Ann Arbor, MI 48409, USA