1. Leibniz Institute for Zoo and Wildlife Research, Alfred-Kowalke-Strasse 17, 10315 Berlin, Germany
2. ISEM, University of Montpellier, CNRS, EPHE, IRD, Montpellier, 34095 Montpellier, France
3. Institute of Biology, Freie Universität Berlin, Takustrasse 6, 14195 Berlin, Germany