1. Centre for Integrated Genomic Medical Research (CIGMR), Centre for Epidemiology, Institute of Population Health, The University of Manchester, Manchester, UK;
2. Formerly of Westlakes Research Institute*, Westlakes Science and Technology Park, Moor Row, Cumbria, UK;
3. Formerly of Northern Genetics Service, Institute of Genetic Medicine, International Centre for Life, Newcastle University, Newcastle upon Tyne, UK;
4. Public Health England, Centre for Radiation, Chemical and Environmental Hazards, Westlakes Science and Technology Park, Moor Row, Cumbria, UK