1. Department of Physiology, School of Medicine, Ehime University, Shigenobu, Onsen-gun, Ehime, Japan 791-02
2. Central Research Laboratories, School of Medicine, Ehime University, Shigenobu, Onsen-gun, Ehime, Japan 791-02
3. Laboratory of Biology, Hitotsubashi University, Kunitachi, Tokyo, Japan 186