1. CAS Key Laboratory of Special Pathogens and Biosafety, Wuhan Institute of VirologyChinese Academy of Sciences 430071 Wuhan Hubei China
2. Hubei Wildlife RescueResearch and Development Center 430074 Wuhan Hubei China
3. Center for Influenza Research and Early warning (CASCIRE)Chinese Academy of Sciences 100101 Beijing China
4. University of Chinese Academy Sciences 101409 Beijing China