1. School of Chemical Engineering and Light Industry, Guangdong University of Technology, Guangzhou, Guangdong 510006, China
2. Guangdong Laboratory of Chemistry and Fine Chemical Engineering Jieyang Center, Jieyang 515200, China
3. Guangdong Provincial Key Laboratory of Plant Resources Biorefinery, Guangdong University of Technology, Guangzhou, Guangdong 510006, China
4. Institute of Microbiology, Guangdong Academy of Sciences, Guangzhou, Guangdong 510070, China
5. School of Life Science and Engineering, Foshan University, Foshan, Guangdong 528000, China