1. Department of Grain Science and Industry, Kansas State University, Manhattan, Kansas 66506, United States
2. Wheat Marketing Center, 1200 NW Naito Pkwy Unit 230, Portland, Oregon 97209, United States
3. Key Laboratories of Crop Genetics and Physiology of the Jiangsu Province, Plant Functional Genomics of the Ministry of Education, Yangzhou University, Yangzhou 225009, China