1. Center for RNA Research, Institute for Basic Science, Seoul 08826, Korea
2. School of Biological Sciences, Seoul National University, Seoul 08826, Korea
3. Soonchunhyang Institute of Medi-bio Science, Soonchunhyang University, Cheonan-si, Chungcheongnam-do 31151, Korea
4. Integrative Omics, Biological Sciences Division, Pacific Northwest National Laboratory, Richland, Washington 99352, United States