1. Key Laboratory of Etiology and Epidemiology of Emerging Infectious Diseases in Universities of Shandong, Taishan Medical College, Taian, Shandong 271000, People’s Republic of China
2. School of Public Health, Taishan Medical College, Taian, Shandong 271000, People’s Republic of China
3. Global Station for Zoonosis Control, Global Institution for Collaborative Research and Education (GI-CoRE), Hokkaido University, Sapporo 001-0020, Japan
4. National Virus Reference Laboratory, School of Medicine, University College Dublin, Belfield, Dublin 4, Ireland