1. Department of Microbiology, Radboud Institute for Biological and Environmental Sciences, Radboud University, Nijmegen 6525 AJ, The Netherlands
2. Department of Ecosystem and Landscape Dynamics, Institute for Biodiversity and Ecosystem Dynamics (IBED), University of Amsterdam, Amsterdam 1098 XH, The Netherlands
3. Department of Earth Sciences─Geochemistry, Utrecht University, Utrecht 3584 CB, The Netherlands
4. Baltic Sea Centre, Stockholm University, Stockholm 114 18, Sweden