1. College of Marine Life Sciences, Department of Marine Ecology, Ocean University of China, Qingdao 266003, China
2. Marine Bioresource and Environment Research Center, Key Laboratory of Marine Eco-Environmental Science and Technology, The First Institute of Oceanography, Ministry of Natural Resources, Qingdao 266061, China
3. Marine Science and Technology College, Zhejiang Ocean University, Zhoushan 316022, China
4. Laboratory for Marine Ecology and Environmental Science, Qingdao Marine Science and Technology Center, Qingdao 266071, China