1. CAS Key Laboratory of Tropical Marine Bio-Resources and Ecology, RNAM Center for Marine Microbiology, Guangdong Key Laboratory of Marine Materia Medica, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou, Guangdong 510301, China
2. College of Oceanology, University of Chinese Academy of Sciences, Qingdao 266400, China
3. Southern Marine Science and Engineering Guangdong Laboratory, Guangzhou, Guangdong 511458, China