1. College of Veterinary Medicine, Southwest University, Chongqing 400715, People’s Republic of China
2. College of Veterinary Medicine, Yangzhou University, Yangzhou, Jiangsu 225009, People’s Republic of China
3. Jiangsu Co-Innovation Center for Prevention and Control of Important Animal Infectious Diseases and Zoonoses, Yangzhou, Jiangsu 225009, People’s Republic of China
4. Department of Pathology & Laboratory Medicine, Tulane University School of Medicine, New Orleans, Louisiana 70112, United States