1. College of Light Industry and Food Engineering, Guangxi University, 100 Daxue Road, Nanning 530004, China
2. National Key Laboratory of Non-food Biomass Energy Technology, National Engineering Research Center for Non-Food Biorefinery, Guangxi Academy of Sciences, 98 Daling Road, Nanning 530007, China
3. Guangxi Zhuangzu Autonomous Region and the Chinese Academy of Sciences, Guangxi Institute of Botany, Guilin 541006, China