1. College of Bioengineering, Key Laboratory of Shandong Microbial Engineering, State Key Laboratory of Biobased Material and Green Papermaking, Qilu University of Technology, Shandong Academy of Sciences, 3501 Daxue Road, Jinan 250353, China
2. Shandong Sunkeen Biological Company, 6789 Xingfuhe Road, Jining 273517, China