1. Key Laboratory of Biology and Genetic Improvement of Horticultural Crops, Ministry of Agriculture and Rural Affairs/Guangdong Vegetable Engineering and Technology Research Center/Lingnan Guangdong Laboratory of Modern Agriculture, College of Horticulture, South China Agricultural University, Guangzhou 510642, China
2. School of Biology and Agriculture, Shaoguan University, Shaoguan 512005, China
3. State Key Laboratory for Conservation and Utilization of Subtropical Agro-Bioresources, College of Life Sciences, South China Agricultural University, Guangzhou 510642, China