1. School of Biological Sciences, the University of Auckland, Private Bag 92019, Auckland 1142, New Zealand
2. Universal Biology Institute, the University of Tokyo, 7-3-1 Hongo, Bunkyo-ku, Tokyo 113-0033, Japan
3. Institute for Advanced Biosciences, Keio University, Tsuruoka, Yamagata, Japan
4. Faculty of Environment and Information Studies, Keio University, Fujisawa, Japan