1. Movement Ecology Laboratory, Department of Ecology, Evolution and Behavior, Alexander Silberman Institute of Life Sciences, The Hebrew University of Jerusalem, Jerusalem 91904, Israel
2. Museum of Vertebrate Zoology, University of California, Berkeley, CA 94720, USA
3. Department of Integrative Biology, University of California, Berkeley, CA 94720-3114, USA
4. School of Food and Agriculture, University of Maine, Orono, ME 04469, USA
5. Oksky State Reserve, pos. Brykin Bor, Spassky raion, Ryazanskaya oblast 391072, Russia
6. School Mathematical Sciences, University of KwaZulu-Natal, Durban, South Africa