1. The Macaulay Institute, Craigiebuckler, Aberdeen AB15 8QH, UK
2. Biomathematics & Statistics Scotland (BioSS), The Macaulay Institute, Craigiebuckler, Aberdeen AB15 8QH, UK ()
3. Institute of Cell, Animal and Population Biology, University of Edinburgh, West Mains Road, Edinburgh EH9 3JT, UK ()