1. Environment and Sustainability Institute, University of Exeter, Penryn TR10 9FE, UK
2. Department of Animal Ecology, Netherlands Institute of Ecology (NIOO-KNAW), PO Box 50, Wageningen 6700 AB, The Netherlands
3. Leibniz Institute of Freshwater Ecology and Inland Fisheries (IGB), Müggelseedamm 310, Berlin 12587, Germany