1. Odum School of Ecology, College of Veterinary Medicine, University of Georgia, Athens, GA, USA
2. Center for the Ecology of Infectious Disease, College of Veterinary Medicine, University of Georgia, Athens, GA, USA
3. Department of Microbiology and Immunology, Montana State University, Bozeman, MT, USA
4. Department of Infectious Disease, College of Veterinary Medicine, University of Georgia, Athens, GA, USA
5. Department of Virology, University of Helsinki, Helsinki, Finland
6. Center for Infectious Disease Dynamics, Pennsylvania State University, University Park, PA, USA
7. Department of Biology, Pennsylvania State University, University Park, PA, USA