1. State Key Laboratory for Conservation and Utilization of Bio-Resources in Yunnan and The Key Laboratory for Southwest Microbial Diversity of the Ministry of Education, Yunnan University, Kunming 650091, People's Republic of China
2. School of Life Science, Yunnan University, Kunming 650091, People's Republic of China
3. Department of Biology, McMaster University, Hamilton, Ontario, Canada L8S 4K1