1. Institute of Integrative and Comparative Biology, University of LeedsLeeds LS2 9JT, UK
2. Department of Humanities, Social and Political Sciences, ETH Zurich, UNO D 11Universitätstrasse 41, 8092 Zurich, Switzerland
3. Department of Ecology and Evolutionary Biology, Princeton University106a Guyot Hall, Princeton University, Princeton, NJ 08544-1003, USA