1. Department of Basic and Translational Sciences, University of Pennsylvania, Philadelphia, PA 19104-6243, USA
2. Department of Organismal Biology and Anatomy, University of Chicago, Chicago, IL 60637, USA
3. Department of Bioengineering, Imperial College London, London SW7 2AZ, UK
4. Department of Biological Sciences, University of Massachusetts, Lowell, MA 01854, USA
5. UMass Movement Center, University of Massachusetts, Lowell, MA 01854, USA