1. University Herbarium, University of California; 1001 Valley Life Sciences Building #2465 Berkeley, California 94720-2465 U.S.A.
2. Department of Biology, Duke University; Durham, North Carolina 27708-0338 U.S.A.
3. Department of Phanerogamic Botany, Swedish Museum of Natural History; Box 50007 SE-104 05 Stockholm Sweden
4. Albrecht-von-Haller-Institut für Pflanzenwissenschaften, Abteilung Systematische Botanik, Georg-August-Universität, Untere Karspüle 2; 37073 Göttingen Germany
5. Department of Biology, Utah State University; Logan, Utah 84322-5305 U.S.A.