1. Wellcome Trust Centre for Human Genetics, University of Oxford, UK
2. Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, UK
3. Department of Hygiene and Epidemiology, University of Ioannina School of Medicine, Ioannina 45110, Greece.
4. Biomedical Research Institute, Foundation for Research and Technology-Hellas, Ioannina, Greece
5. Centre for Genetic Epidemiology and Modeling, Institute for Clinical Research and Health Policy Studies, Department of Medicine, Tufts University School of Medicine, USA