1. Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Leninskiy pr. 33, 119071 Moscow, Russia
2. Key Laboratory of Animal Ecology and Conservation Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China
3. Krasnodar Department of the Azov-Black Sea Branch of the VNIRO (AzNIIRKh), 350000 Krasnodar, Russia