1. Biosciences Institute, Faculty of Medical Sciences, Newcastle University, Newcastle upon Tyne NE2 4HH, UK
2. Sunderland Eye Infirmary, Queen Alexandra Rd., Sunderland SR2 9H, UK
3. Manchester Royal Eye Hospital, Manchester M13 9WL, UK
4. Gyroscope Therapeutics Limited, a Novartis Company, Rolling Stock Yard, 6th Floor, 188 York Way, London N7 9AS, UK
5. Clinical and Translational Research Institute, Faculty of Medical Sciences, Newcastle University, Newcastle upon Tyne NE2 4HH, UK