1. Key Laboratory of Sichuan Province for Conservation and Utilization of Fishes Resources in the Upper Reaches of the Yangtze River, College of Life Sciences, Neijiang Normal University, Neijiang 641100, China
2. College of Fisheries, Guangdong Ocean University, Zhanjiang 524088, China
3. Animal Nutrition Institute, Sichuan Agricultural University, Chengdu 611130, China