1. Key Laboratory of Animal Breeding Reproduction and Molecular Design for Jiangsu Province, College of Animal Science and Technology, Yangzhou University, Yangzhou 225009, China
2. Institutes of Agricultural Science and Technology Development, Yangzhou University, Yangzhou 225009, China
3. Joint International Research Laboratory of Agriculture and Agri-Product Safety of Ministry of Education of China, Yangzhou University, Yangzhou 225009, China
4. Animal & Avian Sciences, University of Maryland, College Park, MA 20742, USA
5. Poultry Institute, Chinese Academy of Agricultural Sciences/Poultry Institute of Jiangsu, Yangzhou 225003, China
6. College of Biotechnology, Jiangsu University of Science and Technology, Zhenjiang 212100, China