1. School of Environmental Sciences, University of Guelph, Guelph, ON N1G2W1, Canada
2. Centro de Investigaciones en Abejas, CUSUR, Universidad de Guadalajara, Enrique Arreola Silva 883, Zapotlan el Grande 49000, Jalisco, Mexico
3. Departamento de Producción Agricola, CUCSUR, Universidad de Guadalajara, Independencia Nal. 161, Autlan 48900, Jalisco, Mexico
4. Department of Population Medicine, University of Guelph, Guelph, ON N1G2W1, Canada