1. Department of Diagnostic Medicine/Pathobiology, College of Veterinary Medicine, Kansas State University, Manhattan, KS 66506, USA
2. Kansas State Veterinary Diagnostic Laboratory, Kansas State University, Manhattan, KS 66506, USA
3. Zoetis, Kalamazoo, MI 49007, USA
4. Foreign Arthropod-Borne Animal Disease Research Unit, National Bio and Agro-Defense Facility, United States Department of Agriculture, Manhattan, KS 66506, USA