1. Department of Ophthalmology, University of California San Francisco, School of Medicine, San Francisco, CA 941583, USA
2. Department of Biochemistry, Downing Site, Cambridge CB2 1QW, UK
3. Department of Anatomy, University of California, San Francisco, CA 94143, USA
4. Cardiovascular Research Institute, University of California, San Francisco, CA 94158, USA
5. Bakar Aging Research Institute, University of California, San Francisco, CA 94143, USA
6. Institute for Human Genetics, University of California, San Francisco, CA 94143, USA