1. College of Fisheries and Life Sciences, Dalian Ocean University, Dalian 116023, China
2. Key Laboratory of Fishery Resources Development and Utilization in South China Sea, South China Sea Fisheries Research Institute, Ministry of Agriculture and Rural Affairs, Guangzhou 510300, China
3. Key Laboratory of Aquatic Product Processing, Ministry of Agriculture and Rural Affairs, South China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Guangzhou 510300, China
4. Key Laboratory of Efficient Utilization and Processing of Marine Fishery Resources of Hainan Province, Sanya 572426, China
5. Shenzhen Base of South China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Shenzhen 518121, China