1. Institute of Preventive Veterinary Medicine, Austral University of Chile, Valdivia 5090000, Chile
2. Institute of Environmental and Evolutionary Sciences, Austral University of Chile, Valdivia 5090000, Chile
3. Reproductive Physiology, Toronto Zoo, Toronto, ON M1B 5K7, Canada
4. Center for Surveillance and Evolution of Infectious Diseases (CSEID), Austral University of Chile, Valdivia 5090000, Chile