1. CAS Key Laboratory of Marine Ecology and Environmental Sciences, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China
2. CAS Engineering Laboratory for Marine Ranching, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China
3. University of Chinese Academy of Sciences, Beijing 100049, China
4. Department of Biology, Duke University, Durham, NC 27708, USA
5. Department of Molecular Biology, Cellular Biology, and Biochemistry, Brown University, Providence, RI 02912, USA