1. The United Graduate School of Agricultural Sciences, Tottori University, 4-101 Koyamacho Minami, Tottori 680-8553, Japan
2. Graduate School of Agricultural and Life Sciences, The University of Tokyo, 1-1-1 Yayoi, Bunkyo, Tokyo 113-8657, Japan
3. Arid Land Research Center, Tottori University, 1390 Hamasaka, Tottori 680-0001, Japan