1. School of Life Sciences, Liaocheng University, Liaocheng 252059, China
2. Beijing Vegetable Research Center (BVRC), Beijing Academy of Agriculture and Forestry Science (BAAFS), Beijing 100097, China
3. National Engineering Research Center for Vegetables, Beijing 100097, China
4. Key Laboratory of Biology and Genetic Improvement of Horticultural Crops (North China), Ministry of Agriculture, Beijing 100097, China
5. Beijing Key Laboratory of Vegetable Germplasm Improvement, Beijing 100097, China
6. Marine Science Research Institute of Shandong Province, Qingdao 266104, China
7. State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, The Chinese Academy of Sciences, Beijing 100093, China
8. China National Botanical Garden, Beijing 100093, China