1. Department of Ophthalmology, Casey Eye Institute, Oregon Health & Science University, Portland, OR 97208, USA
2. Department of Biomedical Engineering, Oregon Health & Science University, Portland, OR 97208, USA
3. Department Integrative Biosciences, School of Dentistry, Oregon Health & Science University, Portland, OR 97208, USA
4. Department Chemical Physiology & Biochemistry, School of Medicine, Oregon Health & Science University, Portland, OR 97208, USA
5. Department of Ophthalmology, Boston University School of Medicine, Boston, MA 02118, USA
6. Department of Anatomy and Neurobiology, Boston University School of Medicine, Boston, MA 02118, USA