1. CAS and Shandong Province Key Laboratory of Experimental Marine Biology, Center for Ocean Mega-Science, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China
2. Laboratory for Marine Biology and Biotechnology, Laoshan Laboratory, Qingdao 266100, China
3. University of Chinese Academy of Sciences, Beijing 101408, China
4. National and Local Joint Engineering Laboratory of Ecological Mariculture, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China
5. Laboratory for Marine Fisheries Science and Food Production Processes, Laoshan Laboratory, Qingdao 266100, China
6. Key Laboratory of Breeding Biotechnology and Sustainable Aquaculture, Institute of Hydrobiology, Wuhan 430072, China