1. Key Laboratory of Forestry Biotechnology of Hunan Province, Central South University of Forestry and Technology, Changsha 410004, China
2. Yuelushan Laboratory Carbon Sinks Forests Variety Innovation Center, Changsha 410012, China
3. Dongan Shunhuang Mountain National Forest Park, Yongzhou 425000, China
4. Forest Bureau of Xiangxi Autonomous Prefecture, Jishou 416000, China
5. College of Biological, Hunan Normal University, Changsha 410081, China