1. Taronga Institute of Science and Learning, Taronga Conservation Society Australia, Mosman, NSW 2088, Australia
2. Conservation Biology Research Group, School of Environmental and Life Sciences, The University of Newcastle, Newcastle, NSW 2308, Australia
3. Beckman Center for Conservation Research, San Diego Zoo Wildlife Alliance, 15600 San Pasqual Valley Road, Escondido, CA 92025, USA
4. School of Earth, Atmospheric and Life Sciences, University of Wollongong, Wollongong, NSW 2522, Australia
5. School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, NSW 2052, Australia