1. Marine College, Shandong University, Weihai 264209, China
2. State Key Laboratory of Satellite Ocean Environment Dynamics, Second Institute of Oceanography, Ministry of Natural Resources, Hangzhou 310051, China
3. Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou), Guangzhou 511458, China
4. Ocean and Ecosystem Science Division, Fisheries and Oceans Canada, Bedford Institute of Oceanography, Dartmouth, NS B2Y 4A2, Canada
5. Jiangsu Key Laboratory of Marine Bioresources and Environment, Co-Innovation Center of Jiangsu Marine Bio-Industry Technology, Jiangsu Ocean University, Lianyungang 222005, China