1. Laboratory of Marine Biotechnology and Chemistry (LBCM), University Bretagne Sud (UBS), EMR CNRS 6076, IUEM, Campus Tohannic, 56000 Vannes, France
2. Research Unit in Process and Environmental Engineering (URGPGE), Faculty of Sciences, PEI, University of Antananarivo, Antananarivo 101, Madagascar
3. SVT Department, Faculty of Sciences, UBS, Campus Tohannic, 56000 Vannes, France