1. Department of Biochemistry, Cancer Biology, Neuroscience and Pharmacology, School of Graduate Studies, Meharry Medical College, Nashville, TN 37208, USA
2. Department of Neurology, Vanderbilt University Medical Center, Nashville, TN 37232, USA
3. Vanderbilt Brain Institute, Nashville, TN 37232, USA
4. Department of Biological Sciences, Vanderbilt University, Nashville, TN 37235, USA