1. College of Forestry, Northeast Forestry University, Harbin 150040, China
2. Key Laboratory of Sustainable Forest Ecosystem Management-Ministry of Education, Northeast Forestry University, Harbin 150040, China
3. Department of Forestry and Forest Resources Management, Faculty of Natural Resources & Environment, University for Development Studies, Nyankpala Campus, Tamale P.O. Box TL 1882, Ghana
4. College of Life Science, Northeast Forestry University, Harbin 150040, China
5. State Forestry and Grassland Administration Engineering Technology Research Center of Korean Pine, Harbin 150040, China