1. National Institute of Agricultural Botany, New Road, East Malling, West Malling, Kent ME19 6BJ, UK
2. The School of Life and Environmental Sciences, University of Lincoln, Lincoln LN6 7DL, UK
3. John Innes Centre, Norwich Research Park, Norwich NR4 7UH, UK
4. Natural Resources Institute, University of Greenwich, Kent ME4 4TB, UK
5. School of Biosciences, University of Kent, Canterbury CT2 7NZ, UK