1. College of Horticulture and Gardening, Yangtze University, Jingzhou 434025, China
2. Beijing Vegetable Research Center, Beijing Academy of Agricultural and Forestry Sciences, Key Laboratory of Biology and Genetic Improvement of Horticultural Crops (North China), Ministry of Agriculture, Beijing Key Laboratory of Vegetable Germplasm Improvement, National Engineering Research Center for Vegetables, Beijing 100097, China
3. Department of Horticulture, Agricultural College, Shihezi University, Shihezi 832003, China