1. School of Life Sciences, East China Normal University, Shanghai 200241, China
2. Key Laboratory of Tropical Hydrobiology and Biotechnology of Hainan Province, Hainan Aquaculture Breeding Engineering Research Center, School of Marine Biology and Fisheries, Hainan University, Haikou 570228, China
3. Commonwealth Scientific and Industrial Research Organisation (CSIRO), Agriculture and Food, Livestock & Aquaculture Program, Bribie Island Research Centre, Bribie Island, Brisbane, QLD 4507, Australia